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中國(guó)重突天牛族Astathini分類和比較形態(tài)學(xué)研究

發(fā)布時(shí)間:2018-08-20 11:12
【摘要】:重突天牛族Astathini隸屬于鞘翅目Coleoptera天?艭erambycidae溝脛天牛亞科Lamiinae。不少種類是農(nóng)林業(yè)生產(chǎn)中的重要害蟲,幼蟲以鉆蛀為害,成蟲以啃食葉片為害。本論文介紹了重突天牛族的國(guó)內(nèi)外分類研究概況、比較形態(tài)學(xué)研究現(xiàn)狀以及該族昆蟲的經(jīng)濟(jì)意義。從分類學(xué)和比較形態(tài)學(xué)兩個(gè)方面開展研究,主要結(jié)果如下:1分類學(xué)研究系統(tǒng)整理、訂正和記述了中國(guó)重突天牛族昆蟲6屬41種(亞種),新增了12種中文描述、12種雄性外生殖器描述以及14種雌性生殖器描述,補(bǔ)充了9種中文描述;拍攝了5屬31種定名標(biāo)本或模式標(biāo)本整體圖、4屬14種雄性外生殖器圖及4屬14種雌性生殖器圖。檢視了模式標(biāo)本5屬18種(亞種),屬級(jí)模式標(biāo)本檢視率達(dá)83%,種級(jí)模式標(biāo)本檢視率達(dá)44%。并整理形成了我國(guó)重突天牛族昆蟲6屬41種(亞種)的名錄發(fā)現(xiàn)了1個(gè)中國(guó)新紀(jì)錄種:老撾纓額天牛Momisis submonticola Breuning,1968,和1個(gè)老撾新紀(jì)錄種:黃藍(lán)眼天牛Bacchisa (Bacchisa) guerryi (Pic,1911)。2比較形態(tài)學(xué)研究屬級(jí)比較形態(tài)學(xué)研究:選取了柄節(jié)形狀、觸角長(zhǎng)度、前胸中瘤突的隆起程度、后胸腹板突伸至中胸基節(jié)窩的程度、鞘翅形狀、腹部顏色、脛節(jié)顏色、雄性腹部第8節(jié)背板形狀、陽基側(cè)突的大小、中莖基和中莖突的長(zhǎng)度之比、雌性腹部第8節(jié)形狀、產(chǎn)卵桿分叉位置等92個(gè)特征,通過對(duì)中國(guó)重突天牛族5屬的比較形態(tài)學(xué)研究,篩選出屬級(jí)具有分類學(xué)意義的特征主要有觸角柄節(jié)長(zhǎng)度、后胸腹板突伸至中足基節(jié)窩的程度、陽基側(cè)突的長(zhǎng)度、端突的形狀以及交配囊的形狀等13個(gè),其中觸角第2節(jié)形狀、雄性腹部第8節(jié)背板形狀、雄性腹部第8節(jié)腹板形狀、陽基側(cè)突長(zhǎng)度、蓋長(zhǎng)度、雌性腹部第8節(jié)形狀、雌性腹部第8節(jié)骨桿長(zhǎng)度、產(chǎn)卵器分叉位置、端突的大小和形狀及交配囊的形狀10個(gè)特征為首次用于該族分屬的特征。種級(jí)比較形態(tài)學(xué)研究:選取了柄節(jié)形狀、額基部是否具縱脊、前胸中瘤突的隆起程度、后胸腹板顏色、鞘翅形狀、腹部腹面顏色、腿節(jié)顏色、陽基側(cè)突形狀中莖基腹面端部形狀、基腹片突端部形狀、端突形狀等92個(gè)特征,通過對(duì)重突天牛族21種的比較形態(tài)學(xué)研究,發(fā)現(xiàn)在不同屬內(nèi)具有分類學(xué)意義的特征不完全相同,種級(jí)具有分類學(xué)意義的特征主要有觸角基瘤之間是否凹陷、柄節(jié)端區(qū)背面特征、前胸中瘤突隆起程度、前胸中瘤突基部?jī)蓚?cè)是否凹入、鞘翅顏色、鞘翅被毛情況、中莖基和中莖突長(zhǎng)度之比、中莖基腹面端部形狀、陽基側(cè)突長(zhǎng)度、內(nèi)囊端部骨針長(zhǎng)度、端突的大小和形狀等28個(gè),其中額基部是否具縱脊、前胸中瘤突基部?jī)蓚?cè)是否凹入、鞘翅被毛情況、鞘翅長(zhǎng)與肩寬之比、陽基側(cè)突長(zhǎng)度、陽基側(cè)突長(zhǎng)寬比、陽基側(cè)突被毛情況、蓋長(zhǎng)度、環(huán)末端形狀、中莖基腹面端部形狀、中莖基與中莖突長(zhǎng)度之比、內(nèi)囊端部骨針長(zhǎng)度、基腹片突端部形狀及端突的大小和形狀14個(gè)特征為首次用于該族分種的特征。比較形態(tài)學(xué)的結(jié)果也顯示生殖器特征,尤其是雄性外生殖器特征在該族種間具有重要的分類學(xué)意義。眼天牛屬Bacchisa種級(jí)具有分類學(xué)意義的特征有柄節(jié)形狀、前胸中瘤突隆起程度、陽基側(cè)突長(zhǎng)度、中莖基腹面端部形狀、中莖基和中莖突長(zhǎng)度之比等17個(gè)。廣翅天牛屬Plaxomicrus種級(jí)具有分類學(xué)意義的特征有觸角顏色、后胸腹板顏色、鞘翅顏色、腹部腹面顏色等9個(gè)。亞重突天牛屬Tetraophthalmus種級(jí)具有分類學(xué)意義的特征有額基部是否具縱脊、前胸中瘤突隆起程度、陽基側(cè)突的長(zhǎng)度、中莖基腹面端部形狀、端突的大小和形狀等14個(gè)。雌雄鑒別特征:雌雄鑒別特征在不同屬間基本相同,主要有觸角長(zhǎng)度:雄蟲觸角長(zhǎng)于雌蟲;腹部第5節(jié)(腹部可見最后1節(jié)):雄性腹部第5節(jié)較小,較平,中央不具細(xì)縱溝;雌性腹部第5節(jié)較大,端部中央微凹,中央具細(xì)縱溝。根據(jù)比較形態(tài)學(xué)研究的結(jié)果,對(duì)黃角梨眼天牛B. fortunei flavicornis和云南亞重突天牛Tetraophthalmus janthinipennis yunnanensis Breuning,1956的分類地位進(jìn)行了訂正,分別為梨眼天牛B. fortunei fortunei和紫翅亞重突天牛T. janthinipennisjanthinipennis的異名。同時(shí)重新編制了中國(guó)重突天牛族分屬、分種檢索表,補(bǔ)充和完善了分類描述。
[Abstract]:Astathini belongs to the subfamily Lamiinae, Cerambycidae, Coleoptera. Many species of larvae are important pests in agricultural and forestry production. The larvae are borers and the adults are leaf-eating. This paper introduces the general situation of taxonomic studies, comparative morphological studies and the present situation of this species in China and abroad. The main results are as follows: 1. Systematic study on Taxonomy and comparative morphology has been carried out to revise and record 41 species (subspecies) of 6 genera of Chinese Cephalotaxus, 12 Chinese descriptions have been added, 12 descriptions of male external genitalia and 14 descriptions of female genitalia have been added to 9 species. Five genera and 18 species (subspecies) of type specimens were inspected. The inspection rate of genus type specimens was 83% and species type specimens was 44%. A new record from China was found in the catalogue: Momisis submonticola Breuning, 1968, and a new record from Laos: Bacchisa guerryi (Pic, 1911). Ninety-two characteristics of the male abdomen, including the shape of sheath and wing, the color of abdomen, the color of tibia, the shape of the back of the eighth abdomen, the size of the lateral process of the penis, the ratio of the length of the middle stem base to the middle stem process, the shape of the eighth abdomen of the female, and the bifurcation position of the ovipositor, were studied. The taxonomic characteristics of the genus were the length of the antennal stalk segment, the extent of the posterior thoracoabdominal protrusion extending to the fossa of the midfoot basal ganglion, the length of the lateral process, the shape of the terminal process and the shape of the mating sac. Length, shape of the 8th segment of the female abdomen, length of the 8th segment of the female abdomen, bifurcation position of the ovipositor, size and shape of the terminal process, and shape of the mating sac are the 10 characteristics first used in the genus. There are 92 characteristics in the shape of basal ventral end, basal ventral end, basal ventral end, basal ventral end, and terminal end in the shape of basal lateral processes. The comparative morphological study of 21 species of Cephalopoda shows that the taxonomic characteristics of the two genera are not identical. The main characteristics of the meaning are whether the antennal basal tumors are concave or not, the dorsal surface of the stalk node area, the degree of the protuberance in the anterior chest, the concave basal part of the protuberance in the anterior chest, the color of the sheath wing, the hair of the sheath wing, the ratio of the length of the middle stem base to the middle stem process, the shape of the ventral end of the middle stem base, the length of the lateral process of the Yang base, the length of the bone needle at the end of the inner Among them, 28 were in shape, including whether the frontal base had a longitudinal ridge, whether the basal part of the midrib was concave, whether the basal part of the midrib was concave, whether the basal part of the midrib was concave, whether the basal part of the midrib was hairy, whether the sheath wing was hairy, whether the ratio of the length of the sheath wing to the width of the shoulder, whether the length of the basal lateral process, whether the basal side process was hairy, whether the cap length of the basal side process, whether The length of the bone needle, the shape of the end of the basal abdominal process, and the size and shape of the end process are the first characteristics of the family. The comparative morphological results also show that the genital characteristics, especially the male external genital characteristics, are of great taxonomic significance among the species. The taxonomic characteristics of Bacchisa species are of great significance. Seventeen were found in the shape of stalk node, the degree of protuberance in the anterior chest, the length of the lateral process in the Yang base, the ventral end shape of the middle stem base, the ratio of the length of the middle stem base to the length of the middle stem process. The taxonomic characteristics of the species were as follows: whether the frontal base had a longitudinal ridge, the degree of tumor protuberance in the anterior chest, the length of the lateral process of the male base, the shape of the ventral end of the middle stem base, the size and shape of the terminal process. The fifth segment of the male abdomen is relatively small and flat with no longitudinal groove in the center; the fifth segment of the female abdomen is larger, with a concave central end and a narrow longitudinal groove in the center. The taxonomic status of SIS Breuning, 1956, was revised as the synonyms of B. fortunei fortunei and T. janthinipennis janthinipennis, respectively.
【學(xué)位授予單位】:西南大學(xué)
【學(xué)位級(jí)別】:碩士
【學(xué)位授予年份】:2015
【分類號(hào)】:S433.5

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